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Eukaryota
  Fungi
    Dikarya
      Ascomycota
            Eurotiomycetes
                    Pyrenulales
                            Pyrenulaceae
                                     Anthracothecium
                                             Anthracothecium biferum
                                             Anthracothecium duplicans
                                             Anthracothecium thwaitesii
                                             Anthracothecium variolosum
                                     Granulopyrenis
                                             Granulopyrenis macrocarpoides
                                                   [Microthelia macrocarpoides]
                                     Pleurotrema
                                             Pleurotrema trichosporum
                                             Pleurotrema uniseriale
                                                   [Anisomeridium uniseriale]
                                                   [Microthelia uniserialis]
                                     Pyrenowilmsia
                                             Pyrenowilmsia ferruginosa
                                                   [Porina ferruginosa]
                                     Pyrenula
                                             Pyrenula aggregata
                                             Pyrenula aspistea
                                             Pyrenula cinerea
                                             Pyrenula knightiana
                                             Pyrenula laevigata
                                               Pyrenula laevigata var. incusa
                                             Pyrenula mamillana
                                                   [Pyrenula marginata]
                                             Pyrenula mastophora
                                             Pyrenula nitida
                                             Pyrenula nitidella
                                             Pyrenula obtecta
                                             Pyrenula pinguis
                                                   [Pyrenula emergens]
                                                   [Pyrenula pinguis var. emergens]
                                             Pyrenula pyrenuloides
                                                   [Anthracothecium pyrenuloides]
                                             Pyrenula subducta
                                             Pyrenula subglabriuscula
                                               Pyrenula subglabriuscula var. natalensis
                                             Pyrenula transparens
                                             Pyrenula wilmsiana